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Prehistoric Ancestry Breakdown

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Source Encyclopedia

Reference Populations and Ancient Source Notes

These entries reuse the bundled source notes for the references used across the model. Browse them as a quick encyclopedia of the populations, sites, locations, and periods represented in the report framework.

Anatolian Neolithic

2 reference entries

c. 6400–6000 calBCE

Barcin Neolithic

Barcin Hoyuk

Marmara, Turkey

Barcın Höyük is among the earliest Neolithic settlements in northwestern Anatolia, situated on the shore of Lake İznik. Its community practised mixed agropastoral subsistence, and genomic evidence from this site has been central to characterising the Anatolian Neolithic gene pool that spread into Europe.

c. 7500–5800 calBCE

Tepecik-Ciftlik Neolithic

Tepecik-Ciftlik

Nigde, Turkey

Tepecik-Çiftlik is a long-lived Neolithic mound in the Cappadocian region of central Anatolia, with evidence for mixed farming, mudbrick architecture, and intensive obsidian use. Individuals belong to the wider Anatolian Neolithic gene pool closely related to populations involved in the spread of farming into the Aegean and the Near East.

Ancient Americas

2 reference entries

c. 7250–6390 calBCE

Kennewick Man

Kennewick individual

Washington State, USA

One of the oldest sequenced genomes from the Americas, recovered from the banks of the Columbia River. Genomically falls within the Native American clade, tracing ancestry to the initial peopling of the Americas via Beringia.

c. 3960–3714 calBCE (~5090 BP)

Moraes Hunter-Gatherers

Moraes rockshelter

Miracatu, São Paulo, Brazil

The Moraes rockshelter in the Ribeira de Iguape valley of southern São Paulo yielded Middle Holocene hunter-gatherer remains associated with inland Atlantic Forest foraging populations. Individuals belong to the broader Native American genetic lineage derived from the initial peopling of the Americas.

Caucasus Hunter-Gatherer

1 reference entries

c. 7940–7599 calBCE

Kotias Mesolithic

Kotias Klde

Georgia

Kotias Klde is a rock shelter in western Georgia that yielded one of the classic reference genomes for Caucasus Hunter-Gatherer (CHG) ancestry. CHG is distinct from both Anatolian Neolithic and Iranian Neolithic lineages, and later contributed to Early Bronze Age steppe populations, giving it broad downstream presence across Eurasia.

Eastern Hunter-Gatherer

2 reference entries

c. 5660–5535 calBCE

Samara Hunter-Gatherer

Lebyazhinka-4

Samara, Russia

Lebyazhinka-4 is a burial site in the Samara region of the middle Volga representing Neolithic foragers of the Eastern European forest-steppe. Individuals are classified as Eastern Hunter-Gatherers and represent the local forager substrate predating Bronze Age steppe pastoralism.

c. 8325–8227 calBCE

Minino Mesolithic

Minino 1

Vologda Oblast, Russia

Minino 1, near Lake Vozhe in northwestern Russia, yielded Mesolithic skeletal remains representing Eastern Hunter-Gatherers. These individuals carried a mixture of Ancient North Eurasian and Western Hunter-Gatherer related ancestry, and EHG ancestry later contributed substantially to Bronze Age steppe populations.

Levantine Neolithic

2 reference entries

c. 6900–6700 BCE

Ain Ghazal Pre Pottery Neolithic C

Ain Ghazal

Amman, Jordan

Ain Ghazal was one of the largest Pre-Pottery Neolithic settlements in the southern Levant, known for its plaster statues and established farming and herding. Individuals belong primarily to a southern Levantine Neolithic lineage rooted in Natufian-related ancestry, with additional Anatolian Neolithic affinity.

Late Pre-Pottery Neolithic B (c. 7000–6600 BCE)

Baja Late Pre Pottery Neolithic B

Ba'ja

Petra region, Jordan

Ba'ja is a Late PPNB settlement near Petra in southern Jordan, notable for its dense architecture and elaborate burials. Individuals belong to the Levantine Neolithic population structure rooted in Natufian-related ancestry, with additional Anatolian Neolithic affinity reflecting broader Near Eastern interaction networks.

Mesolithic and Neolithic East Asia

3 reference entries

c. 4300–1900 calBCE

Early Jomon

Odake Shell Midden

Toyama, Japan

The Odake shell midden in Toyama Prefecture is associated with the Early Jōmon culture, the pre-agricultural forager tradition of the Japanese archipelago. Jōmon individuals carry ancestry deeply divergent from both continental East Asian and West Eurasian populations, rooted in early Upper Paleolithic dispersals into East Asia.

Neolithic

Amur River Neolithic

Songhua River region

Heilongjiang, China

This sample derives from sites along the Songhua River in northeastern China, representing early ceramic-using populations of the Amur Basin. Individuals carry East Asian ancestry closely related to the ancestral pool of later Tungusic and northern East Asian groups.

Middle Neolithic

Yellow River Middle Neolithic

Wanggou site

Henan, China

The Wanggou site in Henan is associated with the Neolithic farming cultures of the Yellow River Basin, which independently developed millet agriculture. Individuals cluster with other East Asian agricultural groups ancestral to the majority of present-day East Asians.

North and East African Neolithic

2 reference entries

c. 3000–1200 BCE

East Africa Pastoral Neolithic

East Africa Pastoral Neolithic horizon

Kenya (Rift Valley)

These samples represent some of the earliest herding populations in eastern Africa, associated with the Savanna Pastoral Neolithic. Ancestry related to this horizon remains particularly high in parts of East Africa today.

c. 5200–4900 calBCE

Ifri n'Amr ou Moussa Neolithic

Ifri n'Amr ou Moussa

Khemisset region, Morocco

Ifri n'Amr ou Moussa is a cave in northern Morocco yielding some of the earliest Neolithic genomic data from northwestern Africa. Individuals derive almost entirely from local Maghrebi ancestry continuous with earlier North African populations, predating the stronger Levant-related input seen in later North African groups.

Oceania and Southeast Asia

2 reference entries

Modern reference

Great Andaman Islands

Andamanese

Andaman Islands

The Andamanese are indigenous to the Andaman Islands and represent one of the most isolated population histories in Asia, tracing ancestry to an early out-of-Africa dispersal predating major agricultural expansions. They have remained largely unaffected by the Neolithic, Bronze Age, and historical gene flow events that reshaped neighboring continental populations.

Modern reference

Papuan-related

Middle Sepik River communities

New Guinea

Papuans carry some of the deepest-diverging non-African ancestry in the human genome, including contributions from both Neanderthal and Denisovan archaic introgression. They descend from an early dispersal out of Africa largely separate from the expansions that shaped mainland Eurasian populations.

Sub-Saharan Africa

3 reference entries

Modern reference

Nilotic-related

Aweil-related Nilotic reference

South Sudan

The Dinka are a Nilotic-speaking pastoral people of South Sudan carrying deep Nile Valley-rooted ancestry with minimal West Eurasian admixture. They serve as a clean reference for modelling sub-Saharan African ancestry in populations where such a component is present.

c. 2576–2465 calBCE

Mota Hunter-Gatherer

Mota Cave

Gamo Highlands, Ethiopia

Mota Cave in southwestern Ethiopia yielded one of the first high-coverage ancient genomes from sub-Saharan Africa. The individual represents a pre-pastoralist East African forager population predating the back-migration of West Eurasian Neolithic-related ancestry into East Africa.

Modern reference

West Africa-related

Yoruba reference

Nigeria

The Yoruba of southwestern Nigeria represent a core West African ancestry profile rooted in the Niger-Congo agricultural tradition. Yoruba-related ancestry is widely used as a proxy for the West African component that spread across sub-Saharan Africa through the Bantu expansion.

Western Hunter-Gatherer

1 reference entries

c. 6100–5900 calBCE

Loschbour Mesolithic

Loschbour Rock Shelter

Müllerthal, Luxembourg

Loschbour is one of the classic reference genomes for Western Hunter-Gatherer (WHG) ancestry in Europe, recovered from a rock shelter in Luxembourg. WHG ancestry formed a major component of pre-Neolithic European populations and persisted through admixture with incoming Anatolian Neolithic farmers.

Zagros Neolithic

1 reference entries

c. 8200–7600 BCE

Ganj Dareh Neolithic

Ganj Dareh

Kermanshah Province, Iran

Ganj Dareh is one of the earliest Neolithic settlements in the Zagros Mountains of western Iran, associated with early herding of goats and a pastoral-focused economy rather than full agriculture. Genetically, individuals belong to an early Iranian Neolithic lineage distinct from Anatolian Neolithic farmers, Levantine groups, and Caucasus Hunter-Gatherers, representing a separate eastern branch of Near Eastern population structure that later contributed ancestry to populations across West Asia and South Asia.